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Reviewer invitation email - username blank, and points to 404 page

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Description of issue or problem I’m having:
The invitation email for reviewers that is populated and sent by OJS no longer has the username filled in, and goes to a blank page: https://veterinaryevidence.org/index.php/ve/reviewer

Steps I took leading up to the issue:
This was reported by our editorial staff, who inform me that they went through the usual steps to generate and send out a reviewer invitation email.

What I tried to resolve the issue:
Discussed with our editors and looked in these forums, but couldn’t find anything specific.

Application Version - e.g., OJS 3.1.2:
OJS 3.1.2.1

Additional information, such as screenshots and error log messages if applicable:

An extract from the email that the reviewer is sent:

To accept or decline the invitation to review please:

1) Log in to the active submissions' site: https://veterinaryevidence.org/index.php/ve/reviewer
2) Click on the submission title in question
3) Follow Step 1 of the Reviewer Steps to indicate whether or not you will undertake the review
Username - {$reviewerUsername}

Any help would be much appreciated.

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